By Dan Papaj
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A more complete list of Dan Papaj's publications [1978-present], along with biographical information, can be found here.

A.S. Leonard, A. Dornhaus, and D.R.Papaj, 2012. Why are floral signals complex? An outline of functional hypotheses. In, Evolution of Plant-Pollinator Interactions. Patiny, S. ed. Cambridge, UK: Cambridge University Press. In press.

R.L. Kascorowski, A.S. Leonard, A. Dornhaus, and D.R. Papaj. 2012. Floral signal complexity as a possible adaptation to environmental variability: a test using nectar-foraging bumble bees.  Animal Behaviour, online. doi: 10.1016/j.anbehav.2012.01.007

C. Nufio and D.R. Papaj. 2012. Aggregative behavior is not explained by an Allee effect in the walnut infesting fly, Rhagoletis juglandis. Journal of Insect Behavior 25:166–182. doi: 10.1007/s10905-011-9287-3

P. Marek, D. Papaj, J. Yeager, S. Molina and W.Moore. 2011. Bioluminescent aposematism in millipedes. Current Biology 21:R680-R681. doi:10.1016/j.cub.2011.08.012

J. Davis, C. Nufio and D.R. Papaj. 2011. Quality or competition: Why increase resource acceptance in the presence of conspecifics? Behavioral Ecology 22: 730–737. doi:10.1093/beheco/arr042

A.S. Leonard and D.R. Papaj. 2011. “X” marks the spot: The possible benefits of nectar guides to bees and plants. Functional Ecology 25: 1–9. doi: 10.1111/j.1365-2435.2011.01885.x

A.S. Leonard, A. Dornhaus, and D.R. Papaj. 2011. Flowers help bees cope with uncertainty: signal detection and the function of floral complexity. Journal of Experimental Biology 214: 113-121 doi: 10.1242/jeb.047407 pdf of bbe (selected as a Faculty 1000 'must read')

Leonard, A.S., A. Dornhaus, and D.R. Papaj. 2011. Forget-me-not: Complex floral displays, inter-signal interactions, and pollinator cognition. Current Zoology 57: 215−224. pdf of bbe

E.C. Snell-Rood, G. Davidowitz and D.R. Papaj. 2011. Reproductive tradeoffs of learning in a butterfly. Behavioral Ecology 22: 291-302. doi: 10.1093/beheco/arq169 pdf of bbe

L.D. Carsten-Conner, D.R. Papaj, and D.M. O'Brien. 2010. Resource allocation to testes in walnut flies and implications for reproductive strategy. Journal of Insect Physiology 56: 1523–1529. doi:10.1016/j.jinsphys.2010.04.024 pdf of bbe

E.C. Snell-Rood and D.R. Papaj. 2009. Patterns of phenotypic plasticity in common and rare environments: A study of host use and color learning in the cabbage white butterfly, Pieris rapae. The American Naturalist 173: 615–631. doi: 10.1086/597609

E.C. Snell-Rood, W. Gronenberg and D.R. Papaj. 2009. Brain size: A global or induced cost of learning? Brain, Behavior and Evolution 73:111–128. doi: 10.1159/000213647 pdf of bbe

I. Kulahci, A. Dornhaus and D.R. Papaj. 2008. Multimodal signals enhance decision making in foraging bumblebees. Proceedings of the Royal Society of London, Series B 275:797–802. doi: 10.1098/rspb.2007.1176

D.R. Papaj and E.C. Snell-Rood. 2007. Commentary: Memory flies sooner from flies that learn faster. Proceedings of the National Academy of Sciences 104:13539-13540. doi: 10.1073/pnas.0706165104 k

D.R. Papaj, H.S. Mallory, and C.A. Heinz. 2007. Extreme weather change and host selection in the pipevine swallowtail, Battus philenor. Oecologia 152:365-375. doi: 10.1007/s00442-007-0658-6 b

K.L. Prudic, A.K. Skemp, and D.R. Papaj. 2007. Aposematic coloration, luminance contrast, and the benefits of conspicuousness. Behavioral Ecology 18:41-46. doi: 10.1093/beheco/arl046 n

E.C. Snell-Rood and D.R. Papaj. 2006. Learning signals within sensory environments: Does host cue learning in butterflies depend on background? Animal Biology 56:173-192. doi: 10.1163/157075606777304203 v

J. Cnaani, J.D. Thomson and D.R. Papaj. 2006. The effect of reward properties on learning and choice in foraging bumblebees. Ethology 112:278–285. doi: 10.1111/j.1439-0310.2006.01174.x v

B.D. Worden and D.R. Papaj. 2005. Flower choice copying in bumblebees. Biology Letters 1(4):504-507. doi: 10.1098/rsbl.2005.0368 v v

D.R. Papaj and G.M. Newsom. 2005. A within-species warning function for an aposematic signal. Proceedings of the Royal Society of London, Series B 272:2519–2523. doi: 10.1098/rspb.2005.3186 b v

D.R. Papaj. 2005. Ovarian dynamics in relation to host quality in the walnut-infesting fly, Rhagoletis juglandis. Functional Ecology 19:396-404. b

S.K. Lynn, J. Cnaani, and D.R. Papaj. 2005. Peak shift discrimination learning as a mechanism of signal evolution. Evolution 59:1300–1305. doi: 10.1554/04-284 b

B.D. Worden, A.K. Skemp and D.R. Papaj. 2005. Learning in two contexts: the effects of interference and body size in bumblebees. Journal of Experimental Biology 208:2045-2053. v

L.D. Carsten and D.R. Papaj. 2005. Effects of reproductive state and host resource experience on mating decisions in a walnut-infesting fly (Rhagoletis juglandis). Behavioral Ecology 16:528–533. b

E.A. Hebets and D.R. Papaj. 2005. Complex signal function: Developing a framework for testable hypotheses. Behavioral Ecology and Sociobiology 57:197-214. doi: 10.1007/s00265-004-0865-7 b

D.R. Papaj. 2004. In praise of a good colleague: Ronald John Prokopy. Journal of Insect Behavior 17: 569-577. v

C.R. Nufio and D.R. Papaj. 2004. Superparasitism of larval hosts by the walnut fly, Rhagoletis juglandis, and its implications for female and offspring performance. Oecologia 141: 460-467. v

C.R. Nufio and D.R. Papaj. 2004. Host marking behaviour as a quantitative signal of competition in the walnut fly Rhagoletis juglandis. Ecological Entomology 29: 336-344.v

J. Cnaani, J.O. Schmidt and D.R. Papaj. 2003. The effect of octopamine on behavioral responses of free-foraging bumblebees to a change in food source profitability. Naturwissenschaften 90:185-188. v

M.R. Weiss and D.R. Papaj. 2003. Butterfly color learning in two different behavioral contexts: How much can a butterfly keep in mind? Animal Behaviour 65: 425-434. c

D.R. Papaj. 2003. Learning. In Encyclopedia of Insect Learning, V. Resh and R. Carde, eds. Academic Press. New York. c

R. Dukas, R.J. Prokopy, D.R. Papaj and J.J. Duan. 2001. Egg-laying behavior of Mediterranean fruit flies: is social facilitation important? Florida Entomologist 84:665-671.

M. Aluja, F. Díaz-Fleischer, D.R. Papaj, G. Lagunes and J. Sivinski. 2001. Effects of age, diet, female density, and the host resource on egg load in Anastrepha ludens and A. obliqua (Diptera: Tephritidae). Journal of Insect Physiology 47:975-988. c

A.D. Lachmann and D.R. Papaj. 2001. Effect of host stimuli on ovariole development in the walnut fly, Rhagoletis juglandis (Diptera, Tephritidae). Physiological Entomology 26:38-48.

C. Nufio and D.R. Papaj. 2001. Host-marking behavior in phytophagous insects and parasitoids. Entomologia Experimentalis et Applicata 99:273-293. c

C.R. Nufio, D. R. Papaj and H. Alonso-Pimentel. 2000. A field analysis of host fruit utilization by the walnut fly, Rhagoletis juglandis. Environmental Entomology 93:994-1001. c

H. Alonso-Pimentel, H.G. Spangler, R. Rogers and D.R. Papaj. 2000. Acoustic component and social context of the wing display of the walnut fly Rhagoletis juglandis. Journal of Insect Behavior 13:511-524.

D.R. Papaj. 2000. Ovarian dynamics and host use. Annual Review of Entomology 45:423-448. b

M.L. Henneman and D.R. Papaj. 1999. Role of host fruit color pattern in the behavior of Rhagoletis juglandis (Diptera: Tephritidae). Entomologia Experimentalis et Applicata 93:249-258.

H. Alonso-Pimentel and D.R. Papaj. 1999. The interaction between effects of resource presence and sex ratio on copulation duration in a walnut fly, Rhagoletis juglandis (Diptera: Tephritidae). Animal Behaviour 57:1063-1069. c

F. Diaz-Fleischer, D.R. Papaj, R.J. Prokopy, A.L. Norrbom and M. Aluja. 1999. Evolution of fruit fly oviposition behavior. pp. 811-842 in Fruit Flies (Tephritidae): Phylogeny and Evolution of Behavior. M. Aluja and A.L. Norrbom (eds.). CRC Press. Boca Raton, Florida. 944p.

R.J. Prokopy and D.R. Papaj. 1999. Behavior of flies of the genera Rhagoletis, Zonosemata and Carpomya (Trypetinae: Carpomyina). pp. 219-252 in Fruit Flies (Tephritidae): Phylogeny and Evolution of Behavior. M. Aluja and A.L. Norrbom (eds.). CRC Press. Boca Raton, Florida. 944p.

H. Alonso-Pimentel, J.B. Korer, C. Nufio and D.R. Papaj. 1998. Role of color and shape stimuli in host-enhanced oogenesis in the walnut fly, Rhagoletis juglandis. Physiological Entomology 23:97-104. c

D.R. Papaj and R.H. Messing. 1998. Asymmetries in dynamical state as an explanation for resident advantage in contests. Behaviour 135:1013-1030.

L.E.M. Vet, A.G. De Jong, E. Franchi, and D.R. Papaj. 1998. The effect of complete versus incomplete information on odour discrimination in a parasitic wasp. Animal Behaviour 55:1271-1279.

R.A. Allard and D.R. Papaj. 1997. Learning of visual cues by pipevine swallowtail butterflies: A test using artificial leaf models. Journal of Insect Behavior 9:961-967.

D.R. Papaj and H. Alonso-Pimentel. 1997. Why walnut flies superparasitize: time savings as a possible function. Oecologia 109:166-174. c

H. Alonso-Pimentel and D.R. Papaj. 1996. Patterns of egg load and their significance for the distribution of a tephritid fly in time and space. Annals of the Entomological Society of America 89:875-882.

H. Alonso-Pimentel and D.R. Papaj. 1996. Operational sex ratio vs. gender densities as determinants of copulation duration in the walnut fly, Rhagoletis juglandis (Diptera: Tephritidae). Behavioral Ecology and Sociobiology 39:171-180.

D.R. Papaj and R.H. Messing. 1996. Functional shifts in the use of parasitized hosts by a tephritid fly: The role of host quality. Behavioral Ecology 7:235-242.

Papaj, D.R., J.M. García and H. Alonso-Pimentel. 1996. Marking of host fruit by male Rhagoletis boycei Cresson flies (Diptera: Tephritidae) and its effect on egg-laying. Journal of Insect Behavior 9:585-598.











Last modified: 10-Feb-2012
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wislizenii photo by Dan Papaj.